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Twenty years after modern civilization has been destroyed, Joel, a hardened survivor, is hired to smuggle Ellie, a 14-year-old girl, out of an oppressive quarantine zone. What starts as a small job soon becomes a brutal, heartbreaking journey, as they both must traverse the United States and depend on each other for survival.

65 million years ago, the knights of the Ryusoul Tribe were selected to participate in a war campaign against the dictating Druidon Tribe who sought to rule Earth. But the Druidon's war campaign was halted by the meteor that brought the age of the dinosaurs to an end, forcing them to flee into space. Meanwhile, the Kishiryu were placed in a deep sleep as a failsafe should the Druidon return. The chosen knights choose to live in peace while passing their mantle down across generations. In the present day, the Druidon return to reclaim Earth as the current generation of knights, the Ryusoulgers, battle them with the support of Ui Tatsui and her father Naohisa Tatsui.

A near future, where humans have been defeated by the viral parasites named Gastrea. They have been exiled into a small territory and live in despair, side-by-side in terror. In this world trapped in darkness— Rentaro, a boy living near Tokyo and member of the "Civil Security"—an organization specializing in fighting against the Gastrea—is used to accomplishing dangerous tasks. His partner is Enju, a precocious young girl. They fight thanks to their peculiar powers until one day, when they receive a special assignment from the government. This top secret mission is to prevent the destruction of Tokyo... Set in a near future, this thrilling heroic-action story... starts now!